Below and above are spread its branches, nourished by the GUNAS; sense-objects are its buds; and below, in the world of men, stretch forth the roots, originating in action.

adhaḥ = downward; ca = and; ūrdhvaṃ = upward; prasṛtāḥ = extended; tasya = its; śākhāḥ = branches; guṇa = by the modes of material nature; pravṛddhāḥ = developed; viṣaya = sense objects; pravālāḥ = twigs; adhaḥ = downward; ca = and; mūlāni = roots; anusantatāni = extended; karma = to work; anubandhīni = bound; manuṣyaloke = in the world of human society.;